Dromaeosaurus (; ) is a genus of Dromaeosauridae theropod dinosaur that lived during the Late Cretaceous period (middle to late Campanian and Maastrichtian), sometime between 80 and 69 million years ago, in the Canada province of Alberta and the western United States. The type species is Dromaeosaurus albertensis, which was described by William Diller Matthew and Barnum Brown in 1922. Its fossils were unearthed in the Hell Creek Formation, Horseshoe Canyon Formation and Dinosaur Park Formation. Teeth attributed to this genus have been found in the Prince Creek Formation. Dromaeosaurus is the type genus of both Dromaeosauridae and Dromaeosaurinae, which include many genera with similar characteristics to Dromaeosaurus such as possibly its closest relative Dakotaraptor. Dromaeosaurus was heavily built, more so than other dromaeosaurids that are similar in size, like Velociraptor.
The first known Dromaeosaurus remains were discovered by paleontologist Barnum Brown during a 1914 expedition to Red Deer River on behalf of the American Museum of Natural History. The area where these bones were collected is now part of Dinosaur Provincial Park in Alberta, Canada. The find, holotype AMNH 5356, consisted of a partial skull in length, a mandible, two , a first metacarpal and some foot bones. The skull lacked most of the top of the snout. Several other skull fragments, and about thirty isolated teeth, are known from subsequent discoveries in Alberta and Montana.
In 1922 William Diller Matthew and Brown named and described the type species of Dromaeosaurus: Dromaeosaurus albertensis. The generic name is derived from the Greek δρομεύς ( dromeus) meaning 'runner' and σαύρος ( sauros) meaning 'lizard'. The specific name, "albertensis", refers to Alberta.
Another seven species of Dromaeosaurus were named: Dromaeosaurus laevifrons (Cope 1876) Matthew & Brown 1922; Dromaeosaurus cristatus (Cope 1876) Matthew & Brown 1922 ( Troodon); Dromaeosaurus? gracilis (Marsh 1888) Matthew & Brown 1922; Dromaeosaurus explanatus (Cope 1876) Kuhn 1939; Dromaeosaurus minutus (Marsh 1892) Russell 1972 (an alvarezsaurid); Dromaeosaurus falculus (Cope 1876) Olshevsky 1979 and Dromaeosaurus mongoliensis (Barsbold 1983) Paul 1988 ( Adasaurus). Most of them were based on fragmentary material, some belonging to other genera, and far less complete than that of Dromaeosaurus albertensis, and those that haven't been reclassified are considered nomen dubium today. Nevertheless, it has grown apparent that Dromaeosaurus albertensis is even rarer in its habitat than other small theropods, although it was the first dromaeosaurid of which reasonably good cranial material was described.
The genus Chirostenotes was considered to be synonymous with Dromaeosaurus at one point in time.
Dromaeosaurus had a relatively robust skull with a deep snout. Its teeth were rather large and were shaped like a curved cone with a coat of enamel covering the crown. It had only nine teeth in each maxilla. Dromaeosaurus also had a vein at the back of the head, the vena capitis dorsalis, that drained the front neck muscles through two long canals running to the posterior surface of the brain. The Meckelian groove of Dromaeosaurus is rather shallow and does not have much depth.
The exact relationships of Dromaeosaurus are somewhat unclear. Although its rugged build gives it a primitive appearance, it was actually a very specialized animal. In an analysis of the clade Dromaeosaurinae, species such as Utahraptor, Achillobator and Yurgovuchia have been recovered. The genus Dakotaraptor has been classified as the Sister group to Dromaeosaurus, but more recent analysis do not recover such a close relationship.
Below is a cladogram by Senter et al. in 2012. Dromaeosaurus is recovered as the sister taxon to Yurgovuchia, Utahraptor and Achillobator.
The cladogram below follows a 2015 analysis by paleontologists Robert DePalma, David Burnham, Larry Martin, Peter Larson, and Robert Bakker, using updated data from the Theropod Working Group. In this analysis, Dromaeosaurus is classified as the sister taxon of Dakotaraptor.
Paleobiology
Feeding behavior
See also
Footnotes
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